He completed a three-year postdoctoral fellowship with the American Cancer Society at the University of California at San Diego (1988-91).
Bartholomew earned his doctoral degree from the University of California-Davis (1988) and his bachelor's degree from Brigham Young University in Salt Lake City (1982).
He currently serves as chair for the National Scientific Review for Genetic Mechanisms in Cancer committee for the American Cancer Society. He has been a scientific reviewer for the U.S. Army and for Science; Molecular Cell, Molecular and Cellular BiologyMethods; Nucleic Acids Research, and Chemistry and Biology.
He has published nearly 30 journal articles and has been awarded seven grants, totaling more than $4.2 million. He currently has funding from the National Institutes of Health for two grants for the study of regulation of gene expression.